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RPS3

CANCER ATLAS

Gene description

 Ribosomal protein S3

Protein class

 Cancer-related genes, Enzymes, Plasma proteins, Predicted intracellular proteins, Ribosomal proteins

Protein evidence

 Evidence at protein level
Colorectal cancer
Breast cancer
Prostate cancer
Lung cancer
Cervical cancer

STAINING SUMMARY
HPA063339

Tissue

Cancer staining

Protein
expression of
normal tissue
Breast cancer
 
Carcinoid
 
Cervical cancer
 
 
Colorectal cancer
 
 
Endometrial cancer
 
 
Glioma
 
Head and neck cancer
 
 
Liver cancer
 
 
Lung cancer
 
 
Lymphoma
 
 
 

Tissue

Cancer staining

Protein
expression of
normal tissue
Melanoma
 
Ovarian cancer
 
Pancreatic cancer
 
Prostate cancer
 
Renal cancer
 
Skin cancer
 
Stomach cancer
 
 
Testis cancer
 
Thyroid cancer
 
Urothelial cancer
 

Staining summary

 Cancer tissues displayed weak to moderate cytoplasmic staining. Rare case of cervical cancer was strongly positive. A majority of carcinoid and lymphoma cases were negative.

GENE INFORMATION

Gene name

 RPS3 (HGNC Symbol)

Synonyms

 FLJ26283, FLJ27450, MGC87870, S3

Description

 Ribosomal protein S3 (HGNC Symbol)

Entrez gene summary

 Ribosomes, the organelles that catalyze protein synthesis, consist of a small 40S subunit and a large 60S subunit. Together these subunits are composed of 4 RNA species and approximately 80 structurally distinct proteins. This gene encodes a ribosomal protein that is a component of the 40S subunit, where it forms part of the domain where translation is initiated. The protein belongs to the S3P family of ribosomal proteins. Studies of the mouse and rat proteins have demonstrated that the protein has an extraribosomal role as an endonuclease involved in the repair of UV-induced DNA damage. The protein appears to be located in both the cytoplasm and nucleus but not in the nucleolus. Higher levels of expression of this gene in colon adenocarcinomas and adenomatous polyps compared to adjacent normal colonic mucosa have been observed. This gene is co-transcribed with the small nucleolar RNA genes U15A and U15B, which are located in its first and fifth introns, respectively. As is typical for genes encoding ribosomal proteins, there are multiple processed pseudogenes of this gene dispersed through the genome. Multiple alternatively spliced transcript variants encoding different isoforms have been found for this gene. [provided by RefSeq, May 2012]

Chromosome

 11

Cytoband

 q13.4

Chromosome location (bp)

 75399486 - 75422280

Protein evidence

 Evidence at protein level (all genes)

Ensembl

 ENSG00000149273 (version 83.38)

Entrez gene

 6188

UniProt

 P23396 (UniProt - Evidence at protein level)

neXtProt

 NX_P23396

Antibodypedia

 RPS3 antibodies

PROTEIN BROWSER
«
RPS3-001
RPS3-004
RPS3-007
RPS3-008
RPS3-013
RPS3-014
RPS3-016
RPS3-019
RPS3-020
RPS3-021
RPS3-022
»

PROTEIN INFORMATION

Splice variant

UniProt

Protein class

Gene ontology

Length & mass

Signal peptide
(predicted)

Transmembrane regions
(predicted)
RPS3-001
ENSP00000434643
ENST00000531188
P23396 [Direct mapping]
40S ribosomal protein S3
Show all
Enzymes
   ENZYME proteins
   Lyases
Ribosomal proteins
Predicted intracellular proteins
Plasma proteins
Cancer-related genes
   Candidate cancer biomarkers
Protein evidence (Kim et al 2014)
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000184 [nuclear-transcribed mRNA catabolic process, nonsense-mediated decay]
GO:0003677 [DNA binding]
GO:0003684 [damaged DNA binding]
GO:0003723 [RNA binding]
GO:0003729 [mRNA binding]
GO:0003735 [structural constituent of ribosome]
GO:0003906 [DNA-(apurinic or apyrimidinic site) lyase activity]
GO:0004520 [endodeoxyribonuclease activity]
GO:0005515 [protein binding]
GO:0005634 [nucleus]
GO:0005730 [nucleolus]
GO:0005737 [cytoplasm]
GO:0005743 [mitochondrial inner membrane]
GO:0005759 [mitochondrial matrix]
GO:0005829 [cytosol]
GO:0005840 [ribosome]
GO:0005844 [polysome]
GO:0005886 [plasma membrane]
GO:0005925 [focal adhesion]
GO:0006281 [DNA repair]
GO:0006351 [transcription, DNA-templated]
GO:0006412 [translation]
GO:0006413 [translational initiation]
GO:0006414 [translational elongation]
GO:0006415 [translational termination]
GO:0006614 [SRP-dependent cotranslational protein targeting to membrane]
GO:0006974 [cellular response to DNA damage stimulus]
GO:0006979 [response to oxidative stress]
GO:0007059 [chromosome segregation]
GO:0007067 [mitotic nuclear division]
GO:0008017 [microtubule binding]
GO:0008134 [transcription factor binding]
GO:0008534 [oxidized purine nucleobase lesion DNA N-glycosylase activity]
GO:0010467 [gene expression]
GO:0010628 [positive regulation of gene expression]
GO:0015631 [tubulin binding]
GO:0015935 [small ribosomal subunit]
GO:0016020 [membrane]
GO:0016032 [viral process]
GO:0017148 [negative regulation of translation]
GO:0019058 [viral life cycle]
GO:0019083 [viral transcription]
GO:0019899 [enzyme binding]
GO:0019900 [kinase binding]
GO:0019901 [protein kinase binding]
GO:0022627 [cytosolic small ribosomal subunit]
GO:0030529 [ribonucleoprotein complex]
GO:0030544 [Hsp70 protein binding]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0031397 [negative regulation of protein ubiquitination]
GO:0032079 [positive regulation of endodeoxyribonuclease activity]
GO:0032183 [SUMO binding]
GO:0032357 [oxidized purine DNA binding]
GO:0032358 [oxidized pyrimidine DNA binding]
GO:0032587 [ruffle membrane]
GO:0042769 [DNA damage response, detection of DNA damage]
GO:0042981 [regulation of apoptotic process]
GO:0043507 [positive regulation of JUN kinase activity]
GO:0044267 [cellular protein metabolic process]
GO:0044390 [ubiquitin-like protein conjugating enzyme binding]
GO:0044822 [poly(A) RNA binding]
GO:0045738 [negative regulation of DNA repair]
GO:0045739 [positive regulation of DNA repair]
GO:0051018 [protein kinase A binding]
GO:0051059 [NF-kappaB binding]
GO:0051092 [positive regulation of NF-kappaB transcription factor activity]
GO:0051225 [spindle assembly]
GO:0051301 [cell division]
GO:0051536 [iron-sulfur cluster binding]
GO:0051879 [Hsp90 protein binding]
GO:0061481 [response to TNF agonist]
GO:0070062 [extracellular exosome]
GO:0070181 [small ribosomal subunit rRNA binding]
GO:0070301 [cellular response to hydrogen peroxide]
GO:0071159 [NF-kappaB complex]
GO:0072686 [mitotic spindle]
GO:0097100 [supercoiled DNA binding]
GO:1901224 [positive regulation of NIK/NF-kappaB signaling]
GO:1902231 [positive regulation of intrinsic apoptotic signaling pathway in response to DNA damage]
GO:1902546 [positive regulation of DNA N-glycosylase activity]
GO:2001235 [positive regulation of apoptotic signaling pathway]
GO:2001272 [positive regulation of cysteine-type endopeptidase activity involved in execution phase of apoptosis]
Show all
 243 aa
26.7 kDa 		No 0
RPS3-004
ENSP00000436971
ENST00000527446
P23396 [Direct mapping]
40S ribosomal protein S3
Show all
Enzymes
   ENZYME proteins
   Lyases
Ribosomal proteins
Predicted intracellular proteins
Plasma proteins
Cancer-related genes
   Candidate cancer biomarkers
Protein evidence (Kim et al 2014)
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000184 [nuclear-transcribed mRNA catabolic process, nonsense-mediated decay]
GO:0003677 [DNA binding]
GO:0003684 [damaged DNA binding]
GO:0003723 [RNA binding]
GO:0003729 [mRNA binding]
GO:0003735 [structural constituent of ribosome]
GO:0003906 [DNA-(apurinic or apyrimidinic site) lyase activity]
GO:0004520 [endodeoxyribonuclease activity]
GO:0005515 [protein binding]
GO:0005634 [nucleus]
GO:0005730 [nucleolus]
GO:0005737 [cytoplasm]
GO:0005743 [mitochondrial inner membrane]
GO:0005759 [mitochondrial matrix]
GO:0005829 [cytosol]
GO:0005840 [ribosome]
GO:0005844 [polysome]
GO:0005886 [plasma membrane]
GO:0005925 [focal adhesion]
GO:0006281 [DNA repair]
GO:0006351 [transcription, DNA-templated]
GO:0006412 [translation]
GO:0006413 [translational initiation]
GO:0006414 [translational elongation]
GO:0006415 [translational termination]
GO:0006614 [SRP-dependent cotranslational protein targeting to membrane]
GO:0006974 [cellular response to DNA damage stimulus]
GO:0006979 [response to oxidative stress]
GO:0007059 [chromosome segregation]
GO:0007067 [mitotic nuclear division]
GO:0008017 [microtubule binding]
GO:0008134 [transcription factor binding]
GO:0008534 [oxidized purine nucleobase lesion DNA N-glycosylase activity]
GO:0010467 [gene expression]
GO:0010628 [positive regulation of gene expression]
GO:0015631 [tubulin binding]
GO:0015935 [small ribosomal subunit]
GO:0016020 [membrane]
GO:0016032 [viral process]
GO:0017148 [negative regulation of translation]
GO:0019058 [viral life cycle]
GO:0019083 [viral transcription]
GO:0019899 [enzyme binding]
GO:0019900 [kinase binding]
GO:0019901 [protein kinase binding]
GO:0022627 [cytosolic small ribosomal subunit]
GO:0030529 [ribonucleoprotein complex]
GO:0030544 [Hsp70 protein binding]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0031397 [negative regulation of protein ubiquitination]
GO:0032079 [positive regulation of endodeoxyribonuclease activity]
GO:0032183 [SUMO binding]
GO:0032357 [oxidized purine DNA binding]
GO:0032358 [oxidized pyrimidine DNA binding]
GO:0032587 [ruffle membrane]
GO:0042769 [DNA damage response, detection of DNA damage]
GO:0042981 [regulation of apoptotic process]
GO:0043507 [positive regulation of JUN kinase activity]
GO:0044267 [cellular protein metabolic process]
GO:0044390 [ubiquitin-like protein conjugating enzyme binding]
GO:0044822 [poly(A) RNA binding]
GO:0045738 [negative regulation of DNA repair]
GO:0045739 [positive regulation of DNA repair]
GO:0051018 [protein kinase A binding]
GO:0051059 [NF-kappaB binding]
GO:0051092 [positive regulation of NF-kappaB transcription factor activity]
GO:0051225 [spindle assembly]
GO:0051301 [cell division]
GO:0051536 [iron-sulfur cluster binding]
GO:0051879 [Hsp90 protein binding]
GO:0061481 [response to TNF agonist]
GO:0070062 [extracellular exosome]
GO:0070181 [small ribosomal subunit rRNA binding]
GO:0070301 [cellular response to hydrogen peroxide]
GO:0071159 [NF-kappaB complex]
GO:0072686 [mitotic spindle]
GO:0097100 [supercoiled DNA binding]
GO:1901224 [positive regulation of NIK/NF-kappaB signaling]
GO:1902231 [positive regulation of intrinsic apoptotic signaling pathway in response to DNA damage]
GO:1902546 [positive regulation of DNA N-glycosylase activity]
GO:2001235 [positive regulation of apoptotic signaling pathway]
GO:2001272 [positive regulation of cysteine-type endopeptidase activity involved in execution phase of apoptosis]
Show all
 243 aa
26.7 kDa 		No 0
RPS3-007
ENSP00000433155
ENST00000530164
F2Z2S8 [Direct mapping]
40S ribosomal protein S3
Show all
Predicted intracellular proteins
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0003723 [RNA binding]
GO:0006281 [DNA repair]
GO:0008534 [oxidized purine nucleobase lesion DNA N-glycosylase activity]
Show all
 117 aa
13.1 kDa 		No 0
RPS3-008
ENSP00000436556
ENST00000525933
H0YEU2 [Direct mapping]
40S ribosomal protein S3
Show all
Predicted intracellular proteins
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0003723 [RNA binding]
GO:0003735 [structural constituent of ribosome]
GO:0005840 [ribosome]
GO:0006281 [DNA repair]
GO:0006412 [translation]
GO:0008534 [oxidized purine nucleobase lesion DNA N-glycosylase activity]
Show all
 171 aa
18.6 kDa 		No 0
RPS3-013
ENSP00000416745
ENST00000422465
E9PSF4 [Direct mapping]
40S ribosomal protein S3
Show all
Predicted intracellular proteins
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0003735 [structural constituent of ribosome]
GO:0005840 [ribosome]
GO:0006281 [DNA repair]
GO:0006412 [translation]
GO:0008534 [oxidized purine nucleobase lesion DNA N-glycosylase activity]
Show all
 103 aa
11.2 kDa 		No 0
RPS3-014
ENSP00000278572
ENST00000278572
P23396 [Direct mapping]
40S ribosomal protein S3
Show all
Enzymes
   ENZYME proteins
   Lyases
Ribosomal proteins
Predicted intracellular proteins
Plasma proteins
Cancer-related genes
   Candidate cancer biomarkers
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000184 [nuclear-transcribed mRNA catabolic process, nonsense-mediated decay]
GO:0003677 [DNA binding]
GO:0003684 [damaged DNA binding]
GO:0003723 [RNA binding]
GO:0003729 [mRNA binding]
GO:0003735 [structural constituent of ribosome]
GO:0003906 [DNA-(apurinic or apyrimidinic site) lyase activity]
GO:0004520 [endodeoxyribonuclease activity]
GO:0005515 [protein binding]
GO:0005634 [nucleus]
GO:0005730 [nucleolus]
GO:0005737 [cytoplasm]
GO:0005743 [mitochondrial inner membrane]
GO:0005759 [mitochondrial matrix]
GO:0005829 [cytosol]
GO:0005840 [ribosome]
GO:0005844 [polysome]
GO:0005886 [plasma membrane]
GO:0005925 [focal adhesion]
GO:0006281 [DNA repair]
GO:0006351 [transcription, DNA-templated]
GO:0006412 [translation]
GO:0006413 [translational initiation]
GO:0006414 [translational elongation]
GO:0006415 [translational termination]
GO:0006614 [SRP-dependent cotranslational protein targeting to membrane]
GO:0006974 [cellular response to DNA damage stimulus]
GO:0006979 [response to oxidative stress]
GO:0007059 [chromosome segregation]
GO:0007067 [mitotic nuclear division]
GO:0008017 [microtubule binding]
GO:0008134 [transcription factor binding]
GO:0008534 [oxidized purine nucleobase lesion DNA N-glycosylase activity]
GO:0010467 [gene expression]
GO:0010628 [positive regulation of gene expression]
GO:0012505 [endomembrane system]
GO:0015631 [tubulin binding]
GO:0016020 [membrane]
GO:0016032 [viral process]
GO:0017148 [negative regulation of translation]
GO:0019058 [viral life cycle]
GO:0019083 [viral transcription]
GO:0019899 [enzyme binding]
GO:0019900 [kinase binding]
GO:0019901 [protein kinase binding]
GO:0022627 [cytosolic small ribosomal subunit]
GO:0030425 [dendrite]
GO:0030529 [ribonucleoprotein complex]
GO:0030544 [Hsp70 protein binding]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0031397 [negative regulation of protein ubiquitination]
GO:0032079 [positive regulation of endodeoxyribonuclease activity]
GO:0032183 [SUMO binding]
GO:0032357 [oxidized purine DNA binding]
GO:0032358 [oxidized pyrimidine DNA binding]
GO:0032587 [ruffle membrane]
GO:0042769 [DNA damage response, detection of DNA damage]
GO:0042981 [regulation of apoptotic process]
GO:0043507 [positive regulation of JUN kinase activity]
GO:0044267 [cellular protein metabolic process]
GO:0044390 [ubiquitin-like protein conjugating enzyme binding]
GO:0044822 [poly(A) RNA binding]
GO:0045738 [negative regulation of DNA repair]
GO:0045739 [positive regulation of DNA repair]
GO:0051018 [protein kinase A binding]
GO:0051059 [NF-kappaB binding]
GO:0051092 [positive regulation of NF-kappaB transcription factor activity]
GO:0051225 [spindle assembly]
GO:0051301 [cell division]
GO:0051536 [iron-sulfur cluster binding]
GO:0051879 [Hsp90 protein binding]
GO:0061481 [response to TNF agonist]
GO:0070062 [extracellular exosome]
GO:0070181 [small ribosomal subunit rRNA binding]
GO:0070301 [cellular response to hydrogen peroxide]
GO:0071159 [NF-kappaB complex]
GO:0072686 [mitotic spindle]
GO:0097100 [supercoiled DNA binding]
GO:1901224 [positive regulation of NIK/NF-kappaB signaling]
GO:1902231 [positive regulation of intrinsic apoptotic signaling pathway in response to DNA damage]
GO:1902546 [positive regulation of DNA N-glycosylase activity]
GO:1990090 [cellular response to nerve growth factor stimulus]
GO:2001235 [positive regulation of apoptotic signaling pathway]
GO:2001272 [positive regulation of cysteine-type endopeptidase activity involved in execution phase of apoptosis]
Show all
 259 aa
28.5 kDa 		No 0
RPS3-016
ENSP00000436335
ENST00000534440
E9PPU1 [Direct mapping]
40S ribosomal protein S3
Show all
Predicted intracellular proteins
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0003723 [RNA binding]
GO:0006281 [DNA repair]
GO:0008534 [oxidized purine nucleobase lesion DNA N-glycosylase activity]
Show all
 158 aa
17.4 kDa 		No 0
RPS3-019
ENSP00000435566
ENST00000528847
E9PJN9 [Direct mapping]
40S ribosomal protein S3
Show all
Predicted intracellular proteins
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0003735 [structural constituent of ribosome]
GO:0005840 [ribosome]
GO:0006281 [DNA repair]
GO:0006412 [translation]
GO:0008534 [oxidized purine nucleobase lesion DNA N-glycosylase activity]
Show all
 36 aa
3.8 kDa 		No 0
RPS3-020
ENSP00000435457
ENST00000527273
E9PK82 [Direct mapping]
40S ribosomal protein S3
Show all
Predicted intracellular proteins
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0003723 [RNA binding]
GO:0006281 [DNA repair]
GO:0008534 [oxidized purine nucleobase lesion DNA N-glycosylase activity]
Show all
 128 aa
14.3 kDa 		No 0
RPS3-021
ENSP00000433421
ENST00000526608
E9PL09 [Direct mapping]
40S ribosomal protein S3
Show all
Predicted intracellular proteins
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0003723 [RNA binding]
GO:0003735 [structural constituent of ribosome]
GO:0005840 [ribosome]
GO:0006281 [DNA repair]
GO:0006412 [translation]
GO:0008534 [oxidized purine nucleobase lesion DNA N-glycosylase activity]
GO:0015935 [small ribosomal subunit]
Show all
 231 aa
25.4 kDa 		No 0
RPS3-022
ENSP00000433821
ENST00000524851
P23396 [Direct mapping]
40S ribosomal protein S3
Show all
Enzymes
   ENZYME proteins
   Lyases
Ribosomal proteins
Predicted intracellular proteins
Plasma proteins
Cancer-related genes
   Candidate cancer biomarkers
Protein evidence (Ezkurdia et al 2014)
Show all
GO:0000184 [nuclear-transcribed mRNA catabolic process, nonsense-mediated decay]
GO:0003677 [DNA binding]
GO:0003684 [damaged DNA binding]
GO:0003723 [RNA binding]
GO:0003729 [mRNA binding]
GO:0003735 [structural constituent of ribosome]
GO:0003906 [DNA-(apurinic or apyrimidinic site) lyase activity]
GO:0004520 [endodeoxyribonuclease activity]
GO:0005515 [protein binding]
GO:0005634 [nucleus]
GO:0005730 [nucleolus]
GO:0005737 [cytoplasm]
GO:0005743 [mitochondrial inner membrane]
GO:0005759 [mitochondrial matrix]
GO:0005829 [cytosol]
GO:0005840 [ribosome]
GO:0005844 [polysome]
GO:0005886 [plasma membrane]
GO:0005925 [focal adhesion]
GO:0006281 [DNA repair]
GO:0006351 [transcription, DNA-templated]
GO:0006412 [translation]
GO:0006413 [translational initiation]
GO:0006414 [translational elongation]
GO:0006415 [translational termination]
GO:0006614 [SRP-dependent cotranslational protein targeting to membrane]
GO:0006974 [cellular response to DNA damage stimulus]
GO:0006979 [response to oxidative stress]
GO:0007059 [chromosome segregation]
GO:0007067 [mitotic nuclear division]
GO:0008017 [microtubule binding]
GO:0008134 [transcription factor binding]
GO:0008534 [oxidized purine nucleobase lesion DNA N-glycosylase activity]
GO:0010467 [gene expression]
GO:0010628 [positive regulation of gene expression]
GO:0015631 [tubulin binding]
GO:0015935 [small ribosomal subunit]
GO:0016020 [membrane]
GO:0016032 [viral process]
GO:0017148 [negative regulation of translation]
GO:0019058 [viral life cycle]
GO:0019083 [viral transcription]
GO:0019899 [enzyme binding]
GO:0019900 [kinase binding]
GO:0019901 [protein kinase binding]
GO:0022627 [cytosolic small ribosomal subunit]
GO:0030529 [ribonucleoprotein complex]
GO:0030544 [Hsp70 protein binding]
GO:0031116 [positive regulation of microtubule polymerization]
GO:0031397 [negative regulation of protein ubiquitination]
GO:0032079 [positive regulation of endodeoxyribonuclease activity]
GO:0032183 [SUMO binding]
GO:0032357 [oxidized purine DNA binding]
GO:0032358 [oxidized pyrimidine DNA binding]
GO:0032587 [ruffle membrane]
GO:0042769 [DNA damage response, detection of DNA damage]
GO:0042981 [regulation of apoptotic process]
GO:0043507 [positive regulation of JUN kinase activity]
GO:0044267 [cellular protein metabolic process]
GO:0044390 [ubiquitin-like protein conjugating enzyme binding]
GO:0044822 [poly(A) RNA binding]
GO:0045738 [negative regulation of DNA repair]
GO:0045739 [positive regulation of DNA repair]
GO:0051018 [protein kinase A binding]
GO:0051059 [NF-kappaB binding]
GO:0051092 [positive regulation of NF-kappaB transcription factor activity]
GO:0051225 [spindle assembly]
GO:0051301 [cell division]
GO:0051536 [iron-sulfur cluster binding]
GO:0051879 [Hsp90 protein binding]
GO:0061481 [response to TNF agonist]
GO:0070062 [extracellular exosome]
GO:0070181 [small ribosomal subunit rRNA binding]
GO:0070301 [cellular response to hydrogen peroxide]
GO:0071159 [NF-kappaB complex]
GO:0072686 [mitotic spindle]
GO:0097100 [supercoiled DNA binding]
GO:1901224 [positive regulation of NIK/NF-kappaB signaling]
GO:1902231 [positive regulation of intrinsic apoptotic signaling pathway in response to DNA damage]
GO:1902546 [positive regulation of DNA N-glycosylase activity]
GO:2001235 [positive regulation of apoptotic signaling pathway]
GO:2001272 [positive regulation of cysteine-type endopeptidase activity involved in execution phase of apoptosis]
Show all
 243 aa
26.7 kDa 		No 0

Contact

The Project

The Human Protein Atlas

The Human Protein Atlas project is funded
by the Knut & Alice Wallenberg foundation.